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1、Beyond Fluorescence Line Height Making sense of chlorophyll fluorescence,Ru Morrison (Ru.Morrisonunh.edu) Mike Behrenfeld (behrenfmscience.oregonstate.edu),You cant see a whale from space but you can see ,Seeing inside phytoplankton from space,Outline,What is phytoplankton fluorescence? The current
2、fluorescence paradox Fluorescence variability Games of chance Changing the odds 2003, 2008, 2009 What fluorescence can do What fluorescence cant do What we need to do!,What is phytoplankton fluorescence?,Fluorescence Efficiency or quantum yield, f ratio of photons out to in,The Fluorescence Paradox,
3、Two potential and mutually incompatible uses Abundance - need constant fluorescence efficiency Physiology - need variable fluorescence efficiency.,Fluorescence variability games of chance,Fluorescence comes from PSII (mainly) f = chance getting an absorbed photon to PSII and then back out again Vari
4、ability comes from quenching Photochemical quenching (qP) Non-photochemical quenching Antennae quenching (qE) PSII RC downregulation / damage (qI) Changes in proportion of PSII to total absorption,NPQ Dominates in Satellite Derived measurements Energy Dependent Quenching decreases f with increasing
5、irradiance PSII downregulation (photoinhibition) suppresses f at all irradiances,Changing the odds - 2003,from Morrison L&O, 2003,PQ,NPQ,Scalar Irradiance,Quantum Yield of Fluorescence,Changing the odds 2008,Behrenfeld et al., BGD, 2009,Nutrient stress,Behrenfeld et al., BGD, 2009,But Schallenberg e
6、t al. (2008) suggested macronutrient stress increased f with unbalanced growth conditions,Detection of iron stress effects.,Behrenfeld et al., BGD, 2009,Changing the odds - 2009,Gulf of Maine case Study Climatologies (2004-2007),Gulf of Maine case Study Climatologies (2004-2007),Dec,Jan,Gulf of Main
7、e case Study Climatologies (2004-2007),Dec,Jan,Seasonal Climatology,Basin Scale Patterns,Goodwin et al., 2009,Timing of stratification onset and spring bloom,What can Fluorescence do,Give insights into phytoplankton physiology Iron stress Photocompensation / acclimation with growth irradiance that i
8、ncorporates hydrodynamics,What Fluorescence cant do,Abundance apart from geographically and spatially limited studies.,What do we need to do?,Minimize calibration uncertainties in the FLH product Develop operational f product Investigate the heck out of it in-light of these recent insights. With current missions How do you separate nutrient and photoacclimation effects? For upcoming missions (GEOCAPE interesting),